It is well known (to the sort of people to whom such things are well known) that the Scottish engineer Fleeming Jenkin was the first to formalize a toy model of Darwin’s evolutionary theory—with results that were most unfavorable to Darwin: The model predicts that random improvements, even when they confer survival advantages, still tend to disappear over the course of a few generations. This was in 1867.
It seems to be far less well known that Jenkin’s model also predicts that all life on earth dies out after a few generations, which would seem to cast doubt on its assumptions. Jenkin was apparently unaware of this, and so, presumably, was Darwin, who gave considerable credence to the Jenkin model in the final edition of The Origin of Species. This was in 1872.
It seems to be even less well known that the inadequacy of Jenkin’s model was identified in a little-noticed letter to the editor of Nature by the mathematician Arthur Sladen Davis. In that letter, Davis corrected Jenkin’s error and supplied an alternative model that he believed was favorable to Darwin. This was in 1871, but apparently Darwin never heard about it.
And it seems to be known only to me (and now, to the readers of this blog!) that Davis’s model is also flawed, in the opposite direction from Jenkin’s, in that it predicts that any species population must grow without bound following the appearance of a beneficial mutation. And as a result of this, the Davis model undercuts Darwin more than it supports him.
I’ve adjusted Davis’s model much as Davis adjusted Jenkin’s, and gotten a result that could be considered favorable to Darwin. In fact, it’s more or less the result that Davis thought he’d gotten, but hadn’t.
Here comes the more technical part. If this is not your cup of tea, stop reading now. Do come back tomorrow though. I’m not always like this.
Continue reading ‘Jenkin Off’